Speak to Handler Shirsha in the New Combine Foothold at -3676, -782, -202 begin the quest.
Find four more lost Drakkel prowler pups in the Waning Forest and Riven Plateau. You can search for them on track and can only have one following you at a time. You may even find all four in the same spot, but here's where we found them:
-3066, -749, -371
-3395, -757, -319
-3085, -752, -269
-3025, -754, -196
Quest auto-completes when you bring him the fourth pup.